Education and positions


1989 Master of Science, University of Turku

1998 Ph.D., University of Turku

1994-1998 Research Scientist, Department of Medical Biochemistry and Molecular Biology, University of Oulu

1999-2002 Academy of Finland Postdoctoral Researcher, University of Oulu

2000 Docent in Molecular Biology, University of Oulu

2002-2007 Academy of Finland Research Fellow, University of Oulu

2004- Project Leader, Biocenter Oulu

2007- Professor of Molecular Biology, University of Oulu

2008- Vice Chair of the Oulu Center for Cell-Matrix Research

2008-2009 1st Vice Director (Research), Institute of Biomedicine, University of Oulu

2010-2012 Chair, Department of Medical Biochemistry and Molecular Biology, University of Oulu

2010- Scientific Director, Biocenter Oulu, University of Oulu

2012-2017 Deputy Director, Academy of Finland Center of Excellence in Cell-Extracellular Matrix Research


Selected scientific advisory functions


2000- Collaborating scientist, FibroGen Inc., San Francisco, CA, USA

2004- Evaluator of grant applications for e.g. Finnish Cultural Foundation, Swedish Research Council, Wellcome Trust (UK), Medical Research Council (UK), Biotechnology and Biological Sciences Research Council (UK), German Research Foundation, Technology Foundation STW (The Netherlands), National Research Agency of France

2008-2013 Member of the Editorial Board, Journal of Biological Chemistry

2009-2013 Member of the Management Committee and Working Group Leader, COST Action TD0901: Hypoxia sensing, signalling and adaptation

2010-2013 Member of the Research Council, University of Oulu

2010-2013 Member of the Faculty Council, Medical Faculty, University of Oulu

2010- Member of the Board, Biocenter Finland

2010-2015 Member of the Research Council for Health, Academy of Finland, 2nd Vice Chair 2010-2012, 1st Vice Chair 2013-2015

2010-2015 Vice Chair of the Subcommittee for International Joint Projects, Academy of Finland


Viimeksi päivitetty: 28.10.2016