National Doctoral Training

• FinBioNet
• Biocenter Finland
• Doctoral Program in Molecular Medicine (DPMM), University of Eastern Finland
• Human Mortality Project (Argumenta)

International Doctoral Training

• International Graduate School in Molecular Medicine Ulm
• University of Lund
• FinGerDevNet
• Wellcome Trust Centre for Cell-Matrix Research, University of Manchester, UK,
• Spemann Graduate School of Biology and Medicine, Albert-Ludwig University in Freiburg, Germany,
• National Laboratory of Protein Engineering and Plant Genetic Engineering, Peking University, China, 2009-2014

• Sino-Finn Life Science Forum: Network of Finnish-Chinese collaboration, including the following institutes:

Chinese partners:

• Wuhan University
• Shanghai Institutes of Biological Sciences, SIBS
• Shanghai Jiao Tong University

Finnish partners:

• Biocenter Kuopio (BCK), University of Eastern Finland
• Biocenter Oulu (BCO), University of Oulu
• Biocentrum Helsinki (BCH), University of Helsinki
• BioCity Turku, University of Turku & Åbo Akademi University
• Institute of Biomedical Technology (IBT), University of Tampere
• Institute of Biotechnology (BI), University of Helsinki
• Institute for Molecular Medicine Finland FIMM, University of Helsinki